Perona

"Ghost Princess" Perona was the commander of the Wild Zombies and Surprise Zombies of Thriller Bark prior to its collapse. She was one of the Mysterious Four and one of the major antagonists of the Thriller Bark Saga. After the defeat of Gekko Moriah's army and during the timeskip, she begrudgingly helped and cooperated with Roronoa Zoro while in Dracule Mihawk's home on Kuraigana Island. Perona has unusually big, round eyes, and has long, light pink hair which she keeps tied into two pigtails with black and white flower hairpins. She wears a red crown with a black cross on her head, as well as red lipstick and a thick layer of black eyeliner. Her black and white striped tights completely cover her legs, and she wears a mini-skirt and a red and gold belt with a heart belt buckle, as well as red buckled boots. She usually keeps her toes pointing inwards in a pigeon-toed manner, and carried around a red parasol shaped like a cute version of a mini devil prior to her battle with Usopp.

As a child, Perona carried the same parasol and wore the same crown and hairpins, but wore her hair in much thicker pigtails and wore a simple, knee-length red dress with a white wave pattern and dots around its center, as well as smaller red boots. She also carried a ghost doll around on a leash, and was accompanied by a bear named Kumae. When first meeting Hogback ten years prior to her encounter with the Straw Hat Pirates, Perona wore the same crown, but the rest of her outfit was not clearly visible, though she seemed to be wearing a similar dress. After the two year timeskip, Perona has a more mature look than before, wearing a long black strapless dress with lavender frills, red high heeled boots and a big black top hat with white floral decor. She has a pink bat tattoo on her left bicep and a watch on her left wrist, and wears pink nail polish on her fingernails. She still has her signature parasol, and also carries a stuffed animal resembling Kumashi with her. Additionally, she now wears her hair in multiple spiral braids on the back of her head. While reading about the Straw Hats' actions on Dressrosa and resulting new bounties, she wore a collared shirt with short ruffled sleeves and a short dark tie hung loosely around her neck. She wore short frilly skirt a a line of crosses. She had short boots with bows on the front. She also had a large straw hat with a bow and large crown on top of it. Perona is very childish and immature, frequently acting on her own whims. She dislikes people telling her what to do and disobeying her will. Despite this, she is normally loyal to Gekko Moriah and apparently sees him as a way of protection, as she was crying to rejoin him after being sent to Kuraigana Island. After Zoro appeared on Kuraigana as well, she tended to his wounds, displaying a willingness to befriend even those she formerly knew as a foe and demonstrating that she can be kind to others, despite her normally cruel demeanor.

She is very fond of cute things, and will only allow someone to work for her if they are cute. Her dream is to create a kingdom where all the cute things in the world have been turned into zombie animals that obey only her. She dislikes Kumashi because, although he is cute in appearance, his voice is not, and thus she prefers him to just be quiet. Her desire for cute things gives her a slightly childish outlook on some issues. She is somewhat playful during combat, threatening to crush Usopp's heart when she was in her ghost form, which nearly caused Usopp to die from fear, and shortly afterward merely laughing it off when revealing that she could not do so. Perona has katsaridaphobia, a fear of cockroaches, to the point of screaming and crying when Usopp covered her with fake ones. She is also somewhat naive, having watched Usopp seemingly lift a 10-ton hammer without questioning whether or not his feat was genuine, and shortly afterward passing out from fear when he attacked her with it, despite not actually being harmed. She is very confident in her Devil Fruit powers being able to defeat anyone; even when confronted by the powerful and dreaded Shichibukai Bartholomew Kuma, after initially being frozen with fear, Perona quickly regained her confidence and proclaimed that she would defeat him. She is also quite stubborn about using them in combat, having continued to use her Negative Hollows against Usopp even after they proved useless due to his negativity because she believed that his immunity was a fluke. After the timeskip, she has become more mature, although she still carries a teddy bear around, indicating that she is still obsessed with cuteness to some degree. Like many others, Perona has her own unique laugh, which uses the word Horo (e.g. "Horo-horo-horo-horo-horo!"); this is based on the name of her Devil Fruit, the Horo Horo no Mi. Being formerly one of the Mysterious Four, Perona had a number of zombies under her command prior to her defection from Thriller Bark. Her ex-crewmate, Doctor Hogback, acknowledges her value under Moriah's employment. While she seems to recognize that bandages can help a person who is wounded, she seems to lack expertise in medicine; during the process of treating Zoro's wounds, she covered him from head to toe in an overabundance of bandages the first time she found him. However, despite this fault she has common sense or enough knowledge of medical skills, as Zoro recovered from his injuries. She also has at least some knowledge of navigation, demonstrated when she brought Zoro to Sabaody Archipelago all the way from Kuraigana Island.